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Notes/Geography/Ecosystems Under Stress
Notes · GeographyUK · A-Levels

Ecosystems Under Stress

An optional physical topic on the functioning of the biosphere and the strains that human activity places upon it. It covers ecosystem structure, energy flow and nutrient cycling, succession to climatic climax, the global biomes, the measurement and threats to biodiversity, and the sustainable management and conservation of ecosystems.

5 sections·~18 min reading time·3 competencies·Level Standard 3 · Advanced 2

T·0666 / 12
Exam profile
AO1 · Understand ecosystem structure, energy and nutrient cycling, succession, biomes and biodiversityAO2 · Apply understanding to explain ecological change and evaluate conservation and sustainable managementAO3 · Interpret food-web, productivity, succession and biodiversity data and maps
Operators:explainanalyseassessevaluateto what extentdescribe the distributioninterpret

basic level

At AS-Level the focus is on describing ecosystem structure, energy flow and the main biomes.

higher level

The full A-Level requires nutrient cycling, succession, the analysis of biodiversity threats and evaluation of sustainable management.

Depth

Reading depth: In depth

Text

Text size: Standard

Contents · 5 sections▾
  1. Ecosystems Under Stress
    • 01Ecosystems: structure, energy and nutrient cycling◐
    • 02Succession and climatic climax◐
    • 03Biomes and a detailed biome study◐
    • 04Biodiversity and ecosystems under threat●
    • 05Conservation, sustainability and case studies●
§ 01

Ecosystems: structure, energy and nutrient cycling#

●●○StandardLPAQA 7037 3.1.6LPDfE GCE Geography - ecosystems

A food web

Food webGraph, grass (producer) → grasshopper, grass (producer) → rabbit, grasshopper → frog, rabbit → fox, frog → hawk, fox → hawkgrass (producer)grasshopperrabbitfrogfoxhawkeaten byeaten by
Fig. 1Interlinked food chains form a food web; energy passes from producers up through the consumers.

Key points

An ecosystem is a community of living organisms (the biotic component) interacting with each other and with the non-living environment (the abiotic component - climate, soil, water, light) within a defined area. Its organisms are organised into trophic (feeding) levels: producers (green plants) fix energy by photosynthesis; primary consumers (herbivores) eat the producers; secondary and tertiary consumers (carnivores) eat other animals; and decomposers (bacteria and fungi) break down dead matter. Feeding relationships form food chains, and because most organisms have several food sources these interlink into food webs, which are more realistic representations of an ecosystem.
Energy flows through an ecosystem in one direction and is progressively lost. Solar energy is captured by producers as gross primary productivity (GPP); after the plants' own respiration, what remains for growth and for the next level is the net primary productivity (NPP). At each transfer up the food chain, only about ten per cent of the energy becomes biomass in the next level - the rest is lost in respiration, movement, heat and in the parts not eaten or not digested. This inefficiency explains why food chains are short and why the amount of energy and biomass falls sharply at each level, producing an ecological pyramid that narrows towards the top.
Nutrients, unlike energy, are recycled within the ecosystem rather than flowing straight through. The Gersmehl model represents this with three stores - biomass (living matter), litter (dead matter on the surface) and soil - linked by transfers: nutrients pass from biomass to litter as leaves and organisms die, from litter to soil by decomposition, and from soil back to biomass by plant uptake. The stores gain from inputs (precipitation and rock weathering) and lose to outputs (runoff and leaching). The relative size of the stores and speed of the transfers differ between ecosystems and reveal how each functions.
The tropical rainforest and a nutrient-poor environment show the contrast clearly. In a rainforest the biomass store is huge and the transfers rapid: warmth and moisture drive fast growth and fast decomposition, so nutrients are cycled quickly and held mainly in the living vegetation rather than the soil - which is why clearing the forest so quickly exhausts the land. Understanding that energy flows and is lost while nutrients cycle and are conserved is the conceptual foundation of the whole topic.
NPP=GPP−R\text{NPP} = \text{GPP} - RNPP=GPP−R

Net primary productivity

Net primary productivity is the energy fixed by producers (GPP) minus what they use in their own respiration (R); it is the energy available to the rest of the ecosystem.

The Gersmehl nutrient-cycle model

Gersmehl nutrient cycleGraph, biomass (living) → litter (dead surface), litter (dead surface) → soil, soil → biomass (living)biomass (living)litter (deadsurface)soilleaf /organism falldecompositionuptake byroots
Fig. 2Nutrients cycle between the biomass, litter and soil stores; inputs come from rain and weathering, outputs from leaching and runoff.

An ecological pyramid of energy

Pyramid of energypyramid, 4 tiers, Data: producers, primary consumers, secondary consumers, tertiary consumersproducers90000primary consumers9000secondary consumers900tertiary consumers90Energy in kJ per m2 per year (representative)
Fig. 3Energy falls by roughly ninety per cent at each transfer, so the pyramid narrows sharply towards the top.
Worked example

Energy transfer efficiency

Producers in an ecosystem fix 90 000 kJ per square metre per year (NPP), and the primary consumers store 9000 kJ per square metre per year. Calculate the efficiency of energy transfer to the primary consumers and explain the loss.

  1. 01Set up the calculation

    Efficiency = (energy transferred / energy available) x 100 = (9000 / 90 000) x 100.

  2. 02Calculate

    This gives 10 per cent.

    900090000×100=10%\dfrac{9000}{90000} \times 100 = 10\%900009000​×100=10%
  3. 03Explain the loss

    The other 90 per cent is lost as heat from respiration and movement, and in parts of the plants not eaten or not digested - which is why so little energy reaches the next level.

Result: The transfer efficiency is 10 per cent; the remaining energy is lost to respiration, movement and material not eaten or digested.

Exam focus

  • Explain energy flow and the roughly ten per cent transfer efficiency, and why this shapes the ecological pyramid.
  • Use the Gersmehl model to explain how nutrient stores and transfers differ between ecosystems (e.g. rainforest versus a poorer ecosystem).

Typical mistakes

  • Saying nutrients flow straight through the ecosystem like energy - nutrients are recycled between the biomass, litter and soil stores.
  • Confusing GPP (total energy fixed) with NPP (what remains after the producers' respiration).

Active revision

Explain, using the Gersmehl model, why clearing a tropical rainforest quickly exhausts the fertility of the land.

Active recall

Recall the key points — then reveal.

Sources: GCE AS and A level subject content for geography (Department for Education) · AQA A-level Geography 7037 specification (AQA)

§ 02

Succession and climatic climax#

●●○StandardLPAQA 7037 3.1.6LPDfE GCE Geography - succession

A lithosere succession

Lithosere succession (illustrative)Timeline from 0 to 150, 0: bare rock, 8: lichens + mosses (pioneers), 30: grasses + herbs, 70: shrubs, 150: climax woodland, 0–150: primary succession (lithosere)0150yearsprimary succession(lithosere)0bare rock8lichens + mosses(pioneers)30grasses + herbs70shrubs150climax woodland
Fig. 4Illustrative lithosere: each seral stage modifies the environment for the next, from bare rock to climax woodland.

Key points

Succession is the change in the plant and animal community of a place over time, as one group of species modifies the environment and is replaced by another. It proceeds through a series of stages, each called a seral stage or sere, from the first colonisers to a final, stable community. Primary succession begins on a completely new surface with no soil - bare rock, a new sand dune, a lava flow; secondary succession begins where an existing community has been cleared but the soil remains - after a fire or the abandonment of a field - and so proceeds faster because the soil and seeds are already there.
In a primary succession the first colonisers are hardy pioneer species (such as lichens and mosses on bare rock) able to tolerate harsh conditions. As they grow and die, they trap moisture and add organic matter, beginning to form a thin soil. This modified environment allows the next stage - grasses and small herbs - to establish; these in turn build deeper soil and shelter, allowing shrubs, then trees, to colonise. At each stage the community makes the environment less hostile and more suitable for the next, and biodiversity and biomass increase.
The succession continues until it reaches a stable, self-perpetuating community in balance with the climate and soil - the climatic climax community. In the British climate this is typically deciduous woodland; in other climates it is the corresponding biome. A named succession illustrates the process: a lithosere begins on bare rock, a psammosere on sand dunes, and a hydrosere in shallow freshwater, each passing through its characteristic seres to woodland.
Human or physical factors often halt succession before it reaches the natural climax, producing an arrested succession or plagioclimax. Grazing, burning, mowing or trampling repeatedly remove the taller vegetation and prevent the woodland climax from developing; heather moorland maintained by burning and grazing is a classic plagioclimax, as is a mown grassland or grazed chalk downland. Recognising a plagioclimax - and explaining the arresting factor that maintains it - is a frequent and discriminating exam requirement, because it shows that the visible vegetation is not always the natural climax.
Worked example

Explaining a plagioclimax

An area of heather moorland shows no sign of developing into woodland despite the climate being suitable for trees. Explain why, using the idea of a plagioclimax.

  1. 01The natural climax

    The climate would naturally support a climax of woodland, so if left alone the moorland would in time succeed to trees.

  2. 02The arresting factor

    The moorland is regularly burned and grazed (for example for grouse and sheep), which removes tree seedlings and the taller vegetation.

  3. 03The result

    This human management arrests the succession at the heather stage, so the moorland is a plagioclimax - held below its natural woodland climax by the arresting factor.

Result: Regular burning and grazing arrest the succession below its woodland climax, maintaining the heather moorland as a plagioclimax.

Exam focus

  • Sequence a named succession (lithosere, psammosere or hydrosere) through its seral stages to the climatic climax.
  • Identify and explain a plagioclimax and the arresting factor (grazing, burning, mowing) that maintains it.

Typical mistakes

  • Confusing primary succession (new surface, no soil) with secondary succession (soil already present, so faster).
  • Assuming the vegetation seen is always the natural climax - grazing or burning may maintain a plagioclimax.

Active revision

Explain how the pioneer species of a primary succession make the environment suitable for later seral stages.

Active recall

Recall the key points — then reveal.

Sources: GCE AS and A level subject content for geography (Department for Education) · AQA A-level Geography 7037 specification (AQA)

§ 03

Biomes and a detailed biome study#

●●○StandardLPAQA 7037 3.1.6LPDfE GCE Geography - biomes

How climate shapes the tropical rainforest

Rainforest functioningGraph, hot + wet climate (aseasonal) → rapid plant growth, rapid plant growth → highest NPP + biodiversity, rapid plant growth → rapid nutrient cycling, rapid nutrient cycling → nutrients held in biomass, nutrients held in biomass → infertile, easily leached soilshot + wetclimate(aseasonal)rapid plantgrowthhighest NPP +biodiversityrapid nutrientcyclingnutrients heldin biomassinfertile,easily leachedsoilswarmth + waterfastdecompositionquick uptakelittle left insoil
Fig. 5The hot, wet climate drives rapid growth and nutrient cycling, giving high productivity but infertile soils.

Key points

A biome is a large-scale global ecosystem, defined by its climate and its characteristic vegetation, such as tropical rainforest, savanna grassland, hot desert, temperate deciduous forest, boreal forest (taiga) and tundra. Their global distribution is controlled principally by climate - especially temperature and precipitation - which is itself governed by latitude and the atmospheric circulation, so the biomes lie in broad latitudinal belts: rainforests near the equator, deserts near the tropics, and the boreal and tundra biomes towards the poles.
The tropical rainforest is the standard detailed biome study and shows how climate shapes ecosystem functioning. Its hot, wet, aseasonal climate (high temperatures and high rainfall all year) drives extremely rapid plant growth, giving the highest net primary productivity and the greatest biodiversity of any biome. The vegetation is stratified into layers - emergents, canopy, under-canopy, shrub and ground layers - each with distinct light and humidity, and the plants show adaptations such as drip-tips, buttress roots and lianas.
A key feature is the nutrient cycle, which, as the Gersmehl model shows, holds the great majority of the nutrients in the living biomass rather than in the soil. Rapid decomposition and immediate uptake by the dense root mat mean nutrients are recycled quickly and the soils are surprisingly infertile. This makes the rainforest fragile: once the trees are removed, the nutrient store is lost, heavy rain leaches the exposed soil, and the land cannot sustain cultivation for long - the basis of the widely observed failure of large-scale clearance.
Contrasting the rainforest with the savanna sharpens the understanding. The savanna's climate has a marked wet and dry season, so grasses dominate with scattered fire- and drought-adapted trees; its productivity is lower and strongly seasonal, and fire is a natural part of its functioning. Comparing two biomes in terms of their climate, structure, productivity and nutrient cycling - and explaining how the climate drives the differences - is exactly the kind of analytical, linking answer the specification rewards.
Worked example

Explaining rainforest fragility

Explain why, despite its enormous biomass, a tropical rainforest is easily and lastingly damaged when it is cleared for agriculture.

  1. 01Where the nutrients are

    The Gersmehl model shows the nutrients are held mainly in the living biomass, not the soil, and are cycled rapidly through fast decomposition and uptake.

  2. 02Effect of clearance

    Removing the trees removes the main nutrient store and stops the recycling; without the canopy, heavy rain leaches the exposed, already-poor soil.

  3. 03The lasting damage

    The land quickly loses its fertility and cannot sustain cultivation, and because the store was in the lost vegetation, recovery is slow - so the damage is lasting.

Result: The nutrients are in the biomass, not the soil, so clearance removes the nutrient store and exposes poor soil to leaching, causing rapid and lasting loss of fertility.

Exam focus

  • Explain the global distribution of biomes in terms of climate and latitude.
  • For a detailed biome, link its climate to its structure, productivity, nutrient cycling and fragility.

Typical mistakes

  • Assuming rainforest soils are fertile because the vegetation is lush - the nutrients are in the biomass, and the soils are poor and easily leached.
  • Describing a biome's vegetation without linking it to the climate that produces it.

Active revision

Compare the climate, productivity and nutrient cycling of the tropical rainforest and the savanna, and explain how climate accounts for the differences.

Active recall

Recall the key points — then reveal.

Sources: GCE AS and A level subject content for geography (Department for Education) · AQA A-level Geography 7037 specification (AQA)

§ 04

Biodiversity and ecosystems under threat#

●●●AdvancedLPAQA 7037 3.1.6LPDfE GCE Geography - biodiversity

Threats to biodiversity

Biodiversity under threatGraph, habitat loss / deforestation → biodiversity loss, pollution → biodiversity loss, over-exploitation → biodiversity loss, invasive species → biodiversity loss, climate change → biodiversity loss, biodiversity loss → reduced resilience + ecosystem serviceshabitat loss /deforestationpollutionover-exploitationinvasive speciesclimate changebiodiversitylossreducedresilience +ecosystem servi…outcompetenativesconditionsshiftfood websdisrupted
Fig. 6Multiple, interacting human pressures reduce biodiversity and degrade the functioning of ecosystems.

Key points

Biodiversity is the variety of life - the number and variety of species, the genetic variation within them, and the range of ecosystems they form. It is valued both for its own sake and for the services it provides: food, medicines, materials, the regulation of climate and water, pollination and soil formation, and cultural and recreational value. Biodiversity is not spread evenly: it is greatest in the tropics, especially the rainforests and coral reefs, and certain biodiversity hotspots contain exceptional concentrations of species found nowhere else, which makes them conservation priorities.
Human activity is placing ecosystems under increasing stress and driving a rapid loss of biodiversity. The main threats are habitat loss and fragmentation (above all deforestation and the conversion of land to agriculture and settlement), pollution (of air, water and land), over-exploitation (overfishing, hunting and harvesting faster than populations can recover), the introduction of invasive alien species that outcompete natives, and climate change, which shifts the conditions faster than many species can adapt or migrate. These threats often act together and reinforce one another.
The consequences reach beyond the loss of individual species. Removing or reducing species can disrupt food webs and the functioning of the whole ecosystem, reduce its resilience and its ability to recover from disturbance, and diminish the ecosystem services on which people depend. Because biodiversity underpins ecosystem stability, its loss can push a stressed ecosystem past a threshold into a degraded state - a coral reef bleaching and dying, or a forest converting to grassland - that is difficult or impossible to reverse.
Measuring and monitoring biodiversity is essential for its management, and geographers use indices of species richness and diversity, and comparisons over time and between places, to assess the health of ecosystems and the effect of threats. The analytical skill the specification develops is to trace how a specific human pressure propagates through an ecosystem to reduce its biodiversity and function, and to evaluate the seriousness of the loss - recognising that some ecosystems, such as coral reefs and tropical rainforests, are both the richest and among the most threatened.

Biodiversity by biome

Relative species richness by biome (illustrative)Column chart: relative species richness by biome, Data: relative richness index · rainforest: 100; relative richness index · coral reef: 92; relative richness index · temperate forest: 55; relative richness index · grassland: 40; relative richness index · tundra: 20; relative richness index · desert: 15020406080100rainforestcoral reeftemperate f…grasslandtundradesert1009255402015relative species richnessbiome
Fig. 7Illustrative relative species richness: greatest in the tropical rainforest and coral reef, least in tundra and desert.
Worked example

Tracing a pressure through an ecosystem

Explain how the introduction of an invasive predator to an island ecosystem can reduce its biodiversity and disrupt its functioning.

  1. 01The introduction

    An invasive predator arrives with no natural controls (predators, parasites or competitors) and native prey have not evolved defences against it.

  2. 02Direct effect

    The predator eats native species faster than they can reproduce, so their populations crash and some may become extinct - a direct loss of biodiversity.

  3. 03Knock-on effect

    Removing those species disrupts the food web (their prey may boom, their own predators lose food), reducing the ecosystem's stability and the services it provides - so the impact spreads beyond the species directly eaten.

Result: The uncontrolled invasive predator drives native species down or to extinction and disrupts the food web, reducing both biodiversity and ecosystem functioning.

Exam focus

  • Explain the main threats to biodiversity and how they interact, with the value of biodiversity and ecosystem services.
  • Trace how a named human pressure reduces the biodiversity and functioning of an ecosystem, and evaluate the seriousness of the loss.

Typical mistakes

  • Listing threats without explaining how they disrupt food webs and reduce ecosystem resilience and services.
  • Treating biodiversity as evenly distributed - it is concentrated in tropical hotspots that are often also the most threatened.

Active revision

'Habitat loss is the most serious threat to global biodiversity.' Assess this statement.

Active recall

Recall the key points — then reveal.

Sources: GCE AS and A level subject content for geography (Department for Education) · AQA A-level Geography 7037 specification (AQA)

§ 05

Conservation, sustainability and case studies#

●●●AdvancedLPAQA 7037 3.1.6LPDfE GCE Geography - conservation

Managing ecosystems for sustainability

Sustainable ecosystem managementGraph, threats + competing players → protected areas, threats + competing players → sustainable use / controlled harvest, threats + competing players → ecotourism (income from conservation), protected areas → conserved, functioning ecosystem, sustainable use / controlled harvest → conserved, functioning ecosystem, ecotourism (income from conservation) → conserved, functioning ecosystem, restoration → conserved, functioning ecosystemthreats +competingplayersprotected areassustainable use/controlledharvestecotourism(income fromconservation)restorationconserved,functioningecosystemrestrictdamagewithincapacitystake forlocalsrebuild
Fig. 8Management balances conservation against the needs of players, using protected areas, sustainable use, ecotourism and restoration.

Key points

Because ecosystems are under such pressure, their management for sustainability - meeting present needs without compromising the ability of future generations to meet theirs - is central to the topic. Management operates at scales from the local (a single nature reserve or a stretch of coast) to the global (international agreements and protected-area networks), and it must reconcile the competing interests of the many players involved: local communities, businesses, governments, conservation groups, scientists and tourists.
A range of strategies is used. Protected areas (national parks, nature reserves, marine protected areas) restrict damaging activities; sustainable use approaches allow controlled harvesting or grazing that does not exceed the ecosystem's capacity to recover; ecotourism aims to generate income from conserving rather than destroying the ecosystem, funding protection and giving local people a stake in it; and restoration seeks to rebuild degraded ecosystems. International frameworks and NGOs coordinate action across borders for migratory species and shared ecosystems.
A local ecosystem study grounds these ideas. A UK example such as a sand-dune system or a heathland shows the interplay of natural succession, human pressure (trampling, development, fire) and management (fencing, boardwalks, controlled grazing, planting). The management aims to conserve the ecosystem and its species while allowing access, and evaluating how well it balances these aims - and the trade-offs involved - is exactly the assessment task, illustrating the general principles at a scale students can investigate directly.
A global case study of a threatened ecosystem, such as a coral reef or an area of tropical rainforest, shows the same tensions at a larger scale and with higher stakes. Coral reefs, for example, are stressed by warming and acidifying oceans, pollution, destructive fishing and tourism, and their management involves marine protected areas, fishing controls and tackling the global driver of climate change - which no local action alone can solve. The evaluative judgement the specification rewards is whether management can genuinely deliver sustainability given the scale of the threats and the competing interests, reaching a supported conclusion rather than an unqualified verdict.
Worked example

Evaluating the management of a coral reef

A coral reef is protected by a marine protected area with fishing controls, yet it continues to decline. Evaluate why the local management may not be enough.

  1. 01What local management achieves

    The marine protected area and fishing controls reduce local pressures - destructive fishing, pollution and physical damage - which genuinely helps the reef's resilience.

  2. 02The global driver

    The reef is also stressed by warming and acidifying oceans caused by global greenhouse-gas emissions, which cause coral bleaching and which no local rule can control.

  3. 03Reach a judgement

    A defensible conclusion is that local management is necessary and buys resilience, but it cannot succeed alone against a global driver - so sustainability requires local action combined with global emissions reduction.

Result: Local protection reduces local stresses but cannot address the global driver of warming, so the reef's sustainability depends on combining local management with global climate action.

Exam focus

  • Explain a range of conservation strategies (protected areas, sustainable use, ecotourism, restoration) and the players involved.
  • Evaluate the effectiveness of management in a named ecosystem, reaching a supported, scale-aware judgement.

Typical mistakes

  • Treating protection as always successful - it must reconcile competing players and, for global drivers like climate change, local action alone is not enough.
  • Describing a case study instead of evaluating how well its management balances conservation and use.

Active revision

'The sustainable management of a threatened ecosystem is impossible while global pressures such as climate change continue.' To what extent do you agree?

Active recall

Recall the key points — then reveal.

Sources: GCE AS and A level subject content for geography (Department for Education) · AQA A-level Geography 7037 specification (AQA)

Contents

Section -- / 05

    • 01Ecosystems: structure, energy and nutrient cycling◐
    • 02Succession and climatic climax◐
    • 03Biomes and a detailed biome study◐
    • 04Biodiversity and ecosystems under threat●
    • 05Conservation, sustainability and case studies●

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Ecosystems Under Stress

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~18
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Competencies
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References & sources

Sources

Department for Education

  • GCE AS and A level subject content for geography

AQA

  • AQA A-level Geography 7037 specification

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